Identalis subsp. matris-occidentalis A habit B inflorescence C sheath, ligule, and blade D subsp. mohinorensis Soreng P.M. Peterson D sheath, ligule, and blade. Drawings from Peterson et al. (2006), A drawn from holotype PD98059 site collection (Peterson 19145 S chez-Alvarado) D drawn from holotype collection (Nesom 6475 McDonald).Revision of Poa L. (Poaceae, Pooideae, Poeae, Poinae) in Mexico: …Figure 11. Poa matris-occidentalis P.M. Peterson Soreng. A, B P. matris-occidentalis subsp. mohinorensis Soreng P.M. Peterson A spikelet B floret; C subsp. matris-occidentalis C spikelet D floret E lower glume F upper glume G floret H palea dorsal view I palea ventral view J perfect flower, stamens, ovary, and lodicules, all enclosed in palea K lodicules L young caryopsis. Drawings from Peterson et al. (2006), A, B drawn from holotype collection (Nesom 6475 McDonald) C drawn from holotype collection (Peterson 19145 S chez-Alvarado).Robert J. Soreng Paul M. Peterson / PhytoKeys 15: 1?04 (2012)scabrous, prickles of moderate coarseness; longest branches 5.5?0 cm, with 3?5 spikelets in the distal 1/3?/2. Spikelets 4? mm long, 1.8?.7 mm wide, broadly lanceolate, laterally compressed, not bulbiferous, greenish to stramineous; florets 2?, hermaphroditic; rachilla internodes terete, 1? mm long, usually hidden, smooth, glabrous; glumes lanceolate, sub-lustrous, equal to subequal, distinctly keeled, keels scabrous distally, upper surfaces often lightly scabrous, edges smooth or lightly scabrous, apex narrowly acute, lower glumes 3? mm long, (1?3-veined (laterals often short), narrowly lanceolate; upper glumes 3.7?.6 mm long, distinctly 3-veined, lanceolate to oblanceolate; calluses dorsally webbed, web distinct, hairs 2? mm long, woolly; lemmas 4.6?.3 mm long, lanceolate, 5-veined, green, distinctly keeled, keel and marginal veins glabrous or sometimes proximally sparsely puberulent, distally scabrous, between veins, muriculate to densely scabrous from near the base, intermediate veins distinct, upper margins narrowly scarious-hyaline, edges lightly scabrous, apices acute to narrowly acute, sometimes briefly purple and bronze tinged; paleas 4.4? mm long, usually distinctly shorter than the lemma, keels long scabrous for most of the length, between the keels moderately muriculate to short aculeolate. Flowers chasmogamous; lodicules (0.3?0.6?.8 mm long, broadly lanceolate to ovate, with a lateral lobe; anthers 2?.2 mm long, infrequently those of distal flower abortive. Caryopses 2.6? mm long, fusiform in side-view, laterally compressed, subtrigonous in cross-section, light brown to olivaceous, sulcus distinct narrow, hilum 0.2?.25 mm long, oval, grain adherent to the palea. 2n = unknown. Discussion. Originally spelled as Poa “matri-occidentalis”, the epithet is correctly spelled as matris-occidentalis (fide Kanchi Gandhi). The species is endemic to high mountains on the west side of the central Sierra Madre Occidental in southern Chihuahua to southwestern Durango (Peterson et al. 2006), and is only known from two peaks that are over 300 km apart. Specimens have sometimes passed under the name Poa tracyi Vasey, a species of the mountains of Colorado and New Mexico (Soreng and Hatch 1983, Soreng 1985, 2007). DNA data (Gillespie and Soreng, unpublished, from the holotype) supports the species placement within Poa subgen. Poa supersect. Homalopoa, rather than in Poa sect. Sylvestres as originally postulated in Peterson et al. (2006) based on order P144 tenuous mor.Identalis subsp. matris-occidentalis A habit B inflorescence C sheath, ligule, and blade D subsp. mohinorensis Soreng P.M. Peterson D sheath, ligule, and blade. Drawings from Peterson et al. (2006), A drawn from holotype collection (Peterson 19145 S chez-Alvarado) D drawn from holotype collection (Nesom 6475 McDonald).Revision of Poa L. (Poaceae, Pooideae, Poeae, Poinae) in Mexico: …Figure 11. Poa matris-occidentalis P.M. Peterson Soreng. A, B P. matris-occidentalis subsp. mohinorensis Soreng P.M. Peterson A spikelet B floret; C subsp. matris-occidentalis C spikelet D floret E lower glume F upper glume G floret H palea dorsal view I palea ventral view J perfect flower, stamens, ovary, and lodicules, all enclosed in palea K lodicules L young caryopsis. Drawings from Peterson et al. (2006), A, B drawn from holotype collection (Nesom 6475 McDonald) C drawn from holotype collection (Peterson 19145 S chez-Alvarado).Robert J. Soreng Paul M. Peterson / PhytoKeys 15: 1?04 (2012)scabrous, prickles of moderate coarseness; longest branches 5.5?0 cm, with 3?5 spikelets in the distal 1/3?/2. Spikelets 4? mm long, 1.8?.7 mm wide, broadly lanceolate, laterally compressed, not bulbiferous, greenish to stramineous; florets 2?, hermaphroditic; rachilla internodes terete, 1? mm long, usually hidden, smooth, glabrous; glumes lanceolate, sub-lustrous, equal to subequal, distinctly keeled, keels scabrous distally, upper surfaces often lightly scabrous, edges smooth or lightly scabrous, apex narrowly acute, lower glumes 3? mm long, (1?3-veined (laterals often short), narrowly lanceolate; upper glumes 3.7?.6 mm long, distinctly 3-veined, lanceolate to oblanceolate; calluses dorsally webbed, web distinct, hairs 2? mm long, woolly; lemmas 4.6?.3 mm long, lanceolate, 5-veined, green, distinctly keeled, keel and marginal veins glabrous or sometimes proximally sparsely puberulent, distally scabrous, between veins, muriculate to densely scabrous from near the base, intermediate veins distinct, upper margins narrowly scarious-hyaline, edges lightly scabrous, apices acute to narrowly acute, sometimes briefly purple and bronze tinged; paleas 4.4? mm long, usually distinctly shorter than the lemma, keels long scabrous for most of the length, between the keels moderately muriculate to short aculeolate. Flowers chasmogamous; lodicules (0.3?0.6?.8 mm long, broadly lanceolate to ovate, with a lateral lobe; anthers 2?.2 mm long, infrequently those of distal flower abortive. Caryopses 2.6? mm long, fusiform in side-view, laterally compressed, subtrigonous in cross-section, light brown to olivaceous, sulcus distinct narrow, hilum 0.2?.25 mm long, oval, grain adherent to the palea. 2n = unknown. Discussion. Originally spelled as Poa “matri-occidentalis”, the epithet is correctly spelled as matris-occidentalis (fide Kanchi Gandhi). The species is endemic to high mountains on the west side of the central Sierra Madre Occidental in southern Chihuahua to southwestern Durango (Peterson et al. 2006), and is only known from two peaks that are over 300 km apart. Specimens have sometimes passed under the name Poa tracyi Vasey, a species of the mountains of Colorado and New Mexico (Soreng and Hatch 1983, Soreng 1985, 2007). DNA data (Gillespie and Soreng, unpublished, from the holotype) supports the species placement within Poa subgen. Poa supersect. Homalopoa, rather than in Poa sect. Sylvestres as originally postulated in Peterson et al. (2006) based on tenuous mor.
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